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ao. Univ.-Prof. Mag. Dr. Doris Nagel

T: +43-1-4277-53524

doris.nagel@univie.ac.at

Summer term 2026

280010 VO Paleoecology and Paleobiogeography

300084 SE Bachelorseminar for students of the teacher program Biology

300118 VU Introduction to Paleobiology

300133 VO Evolution

300143 UE Research methods in actuopaleobiology: research and teaching

300212 UE Palaeontological field school

300269 VO Palaeobiogeography

300279 EX Palaeontological excursion - Highlights 2

Winter term 2025

280195 PR Individual Research Project Geology, Paleobiology and Geobiology

300084 SE Bachelorseminar for students of the teacher program Biology

300138 VO The last Ice Age - the world between 2,5 million and 10.000 years before present

300163 PP Form and function of organisms - Participation in the workgroups

300238 VU Introduction to Paleobiology

300390 EX Highlights of palaeontology 1 - excursion

300481 VO Morphology and Palaeodiversity of vertebrates - Mammals and Birds

300483 UE Palaeontological Working Methods - Laboratory

Summer term 2025

280010 VO Paleoecology and Paleobiogeography

280176 VU Actuopaleontology

300084 SE Bachelorseminar for students of the teacher program Biology

300118 VU Introduction to Paleobiology

300133 VO Evolution

300143 UE Research methods in actuopaleobiology: research and teaching

300212 UE Palaeontological field school

300269 VO Palaeobiogeography

300279 EX Palaeontological excursion - Highlights 2

Nagel, D., Kavcik-Graumann, N., Rabeder, G., Alberti, F., Döppes, D., & Lindauer, S. (2024). Jungpleistozäne Säugetierreste aus der Schwaigerhöhle bei Mixnitz im Grazer Bergland. Die Höhle: Zeitschrift für Karst und Höhlenkunde, 75(1-4), 189-202.

Marciszak, A., & Nagel, D. (2024). One or two species? Revision of fossil martens from the late Early Pleistocene sites Deutsch Altenburg 2 and 4 (Austria). Palaeontologia Electronica, 27(2), Article a32. https://doi.org/10.26879/1397

Tsoukala, E., Nagel, D., YOULATOS, DIONISIOS., CRÉGUT-BONNOURE , EVELYNE., & SPASSOV , NIKOLAI. (2024). PRIMATES AND CARNIVORES FROM LATE MIOCENE AND EARLIEST LATE PLIOCENE SITES OF MACEDONIA, NORTHERN GREECE. Fossil Imprint, 8, 362–389.

Baca, M., Popović, D., Agadzhanyan, A. K., Baca, K., Conard, N. J., Fewlass, H., Filek, T., Golubiński, M., Horáček, I., Knul, M. V., Krajcarz, M., Krokhaleva, M., Lebreton, L., Lemanik, A., Maul, L. C., Nagel, D., Noiret, P., Primault, J., Rekovets, L., ... Nadachowski, A. (2023). Ancient DNA of narrow-headed vole reveal common features of the Late Pleistocene population dynamics in cold-adapted small mammals. Proceedings of the Royal Society of London B: Biological Sciences, 290(1993), Article 20222238. https://doi.org/10.1098/rspb.2022.2238

Nagel, D., Filek, T., & Tuzar, J. M. (2023). Das Tagebuch des Josef Bayer. 150 Jahre Forschungsgeschichte der Fuchsen- oder Teufelslucke (1824-2023). Das Waldviertel: Zeitschrift für Heimat- und Regionalkunde, 72(2), 235-245.

Nagel, D. (2023). Josef Bayer (1882-1931). Tagebuchnotizen zu den Ausgrabungen in der Teufelslucke bei Roggendorf, 1929 und dem Zwergenloch bei Rosenburg, 1928 in Niederösterreich. Publikationen der Krahuletz-Gesellschaft Eggenburg, 2023(4), 1-37. https://doi.org/http://www.krahuletzmuseum.at/wp-content/uploads/2024/01/PubKraGes-2023_04_Bayer-1.pdf

Gimranov, D., Bocherens, H., Kavcik-Graumann, N., Nagel, D., & Rabeder, G. (2023). The cave bears from Imanay Cave (Southern Urals, Russia). Historical Biology, 35(4), 580-588. https://doi.org/10.1080/08912963.2022.2056837

Nagel, D., Hofreiter, M., Alberti, F., Lindauer, S., Kavcik-Graumann, N., & Rabeder, G. (2023). The climate of the Alps of the last 65,000 years. Climate reconstruction according to fossil vertebrate remains.. Paper presented at 28th International Cave Bear Symposium 2023, Bayreuth, Germany.

Nagel, D., Döppes, D., & Rabeder, G. (2023). The elusive Drachenhöhle near Mixnitz (Styria, Austria). How to date an undatable cave. Paper presented at 28th International Cave Bear Symposium 2023, Bayreuth, Germany.

Morlo, M., Hohenegger, J., & Nagel, D. (2022). A new quantitative approach for guild analyses reveals the evolution of the carnivoran (Mammalia) guild at the Upper Miocene Proto-Rhine (Germany). Historical Biology, 34(8), 1337-1346. https://doi.org/10.1080/08912963.2021.1970153

Kullmer, O., Sandrock, O., Viola, T. B., Hujer, W., Bedaso, Z., Nagel, D., Stadlmayer, A., Popp, F., Scholger, R., Weber, G., & Seidler, H. (2022). Miocene to Pliocene stratigraphy and paleoecology of Galili, Ethiopia. In S. C. Reynolds, & R. Bobe (Eds.), African Paleoecology and Human Evolution (pp. 242-255). Cambridge University Press. https://doi.org/10.1017/9781139696470.020

Xafis, A., Mayda, S., Alçiçek, C. M., Kaya, T., Halaçlar, K., Grimsson, F., & Nagel, D. (2021). Large giraffids (Mammalia, Ruminantia) from the new late Miocene fossiliferous locality of Kemiklitepe-E (Western Anatolia, Turkey). Palaeobiodiversity and Palaeoenvironments, 101(3), 853–867. https://doi.org/10.1007/s12549-020-00433-4

Xafis, A., Wolfgring, E., Nagel, D., & Grimsson, F. (2020). The ever-browsing Deinotheriidae (Mammalia, Proboscidea): Did climate change affect conservative herbivory during the Miocene?. 59. Paper presented at 64th Palaeontological Association Annual Meeting, Oxford, United Kingdom.

Morlo, M., Nagel, D., & Bastl, K. A. (2020). Evolution of the carnivoran (Carnivora, Mammalia) guild structure across the Middle/Upper Miocene boundary in Germany. Palaeogeography, Palaeoclimatology, Palaeoecology, 553, Article 109801. https://doi.org/10.1016/j.palaeo.2020.109801

Stanton, D. W. G., Alberti, F., Plotnikov, V., Androsov, S., Grigoriev, S., Fedorov, S., Kosintsev, P., Nagel, D., Vartanyan, S., Barnes, I., Barnett, R., Ersmark, E., Döppes, D., Germonpré, M., Hofreiter, M., Rosendahl, W., Skoglund, P., & Dalén, L. (2020). Early Pleistocene origin and extensive intra‑species diversity of the extinct cave lion. Scientific Reports, 10, Article 12621. https://doi.org/10.1038/s41598-020-69474-1

Xafis, A., Saarinen, J., Bastl, K., Nagel, D., & Grímsson, F. (2020). Palaeodietary traits of large mammals from the middle Miocene of Gračanica (Bugojno Basin, Bosnia-Herzegovina). Palaeobiodiversity and Palaeoenvironments, 100, 457–477. https://doi.org/10.1007/s12549-020-00435-2

Bastl, K. A., Nagel, D., Morlo, M., & Göhlich, U. B. (2020). The Carnivora (Mammalia) from the middle Miocene locality of Gračanica (Bugojno Basin, Gornji Vakuf, Bosnia and Herzegovina). Palaeobiodiversity and Palaeoenvironments, 100(2), 307-319. https://doi.org/10.1007/s12549-018-0353-0

Westbury, M. V., Hartmann, S., Barlow, A., Preick, M., Ridush, B., Nagel, D., Rathgeber, T., Ziegler, R., Baryshnikov, G., Sheng, G., Ludwig, A., Wiesel, I., Dalén, L., Bibi, F., Werdelin, L., Heller, R., & Hofreiter, M. (2020). Hyena paleogenomes reveal a complex evolutionary history of cross-continental gene flow between spotted and cave hyena. Science Advances, 6(11). https://doi.org/10.1126/sciadv.aay0456 Article

Morlo, M., Le Maitre, A., Bastl, K. A., Engel, T., Lutz, H., Lischewsky, B., von Berg, A., & Nagel, D. (2019). First record of the mustelid Trochictis (Carnivora, Mammalia) from the early Late Miocene (MN 9/10) of Germany and a re-appraisal of the genus Trochictis. Historical Biology, 33(8), 1183-1195.

Morlo, M., Miller, E. R., Bastl, K. A., Abdelgawad, M. K., Hamdan, M., & Nagel, D. (2019). New Amphicyonids (Mammalia, Carnivora) from Moghra, Early Miocene, Egypt. Geodiversitas, 41(1), 731-745.

Doris Nagel ao.Univ.Prof. Dr.Mag.
Department of Palaeontology

Postal address:
Althanstrasse 14
1090 Austria

Email: doris.nagel@univie.ac.at
Phone: +43-1-4277-53524

Research interests

Palaeecology of Mammals (Neogene to Pleistocene)

Terrestrial Actuopalaeontology

Conservation of National fossil sites

The evolution and adaption of mammals to various environments since their diversification from the Paleocene on has been a fascination area since the beginning of my studies. Different aspects of these investigations are food preferences specially in carnivorous mammals, evolution of the middle and inner ear, morphological adaption of the locomotion system as well as migration routes.
Each of these topics needs specialised investigation techniques. Micro-wear analyses and high speed cameras give insight into feeding habits, computertomography allows a non invasive look into the secrets of hearing, and ancient DNA allows detailed reconstructions of centre of origin and migration routes

Projects

Alpinet Culture 2000

Nagel, D. (Project Lead)

1/09/04 → 30/06/07

Das Sarmatium von Nexing - Pilotstudie Mikrofauna

Nagel, D. (Project Lead)

1/09/16 → 31/08/17

Die Teufelslucke (Eggenburg, NÖ) - Archiv der Klimaschwankungen im Jungpleistozän Niederösterreichs

18/10/21

1 Media contribution

Publications

Dinosaur Provincial Park

Seyfullah, L. J. (Visiting researcher), Nagel, D. (Visiting researcher) & Halbwidl, K. (Visiting researcher)

29 Jun 2025 → 6 Jul 2025

The elusive Drachenhöhle near Mixnitz (Styria, Austria) – how to date an undatable cave

Nagel, D. (Speaker), Döppes, D. (Speaker) & Rabeder, G. (Speaker)

22 Sept 2023

Als es der Hyäne bei uns zu kalt wurde. 50.000 Jahre Klimageschichte aus der Teufelslucke bei Roggendorf.

Nagel, D. (Speaker)

28 Oct 2022

Die Teufelslucke bei Eggenburg – Klimaarchiv der letzten 50.000 Jahre

Nagel, D. (Speaker)

4 Jul 2022

Pint of Science- tales from the 50 million year old amber forest

Seyfullah, L. J. (Speaker) & Nagel, D. (Speaker)

11 May 2022

How cold and how dry. New data from the Teufelslucke (Lower Austria, MIS 3)

Filek, T. (Speaker), Nagel, D. (Speaker), Döppes, D. (Speaker) & Rabeder, G. (Speaker)

1 Mar 2022 → 31 Mar 2022

Predation on Allactaga major (Teufelslucke, Lower Austria, Upper Pleistocene): carnivores or birds of prey

Filek, T. (Contributor) & Nagel, D. (Contributor)

12 Oct 2019

The Schwaiger cave near Mixnitz (Styria, Austria)

Nagel, D. (Contributor), Döppes, D. (Contributor) & Rabeder, G. (Contributor)

20 Sept 2019

The giraffe that went west: the first record of Alcicephalus neumayri (Mammalia, Ruminantia) from the late Miocene of Anatolia

Xafis, A. (Contributor), Mayda, S. (Contributor), Grimsson, F. (Contributor), Nagel, D. (Contributor), Kaya, T. (Contributor) & Halaçlar, K. (Contributor)

2 Jul 2019

Palaeodiets of large mammals from the Middle Miocene of Gračanica (Bugojno Basin), Bosnia-Herzegovina. A multiproxy approach

Xafis, A. (Speaker), Saarinen, J. (Contributor), Bastl, K. A. (Contributor), Grimsson, F. (Contributor) & Nagel, D. (Contributor)

17 Mar 2019

A basal turtle shows its teeth: possible evidence of dental microwear on the palatal denticles of Proganochelys quenstedti (Testudinata; Late Triassic)

Xafis, A. (Speaker), Sterli, J. (Contributor), Nagel, D. (Contributor) & Vlachos, E. (Contributor)

14 Dec 2018 → 17 Dec 2018

Fossil Giraffidae (Mammalia, Artiodactyla) from the early Turolian of Kavakdere (Central Anatolia, Turkey)

Xafis, A. (Speaker), Mayda, S. (Contributor), Nagel, D. (Contributor), Grimsson, F. (Contributor) & Kaya, T. (Contributor)

14 Dec 2018 → 17 Dec 2018

Palaeodietary traits of large mammals from the early Middle Miocene fossiliferous locality of Gracanica (Bugojna Basin), Bosnia-Herzegovina

Nagel, D. (Contributor), Xafis, A. (Speaker), Bastl, K. A. (Contributor), Grimsson, F. (Contributor) & Saarinen, J. (Contributor)

12 Oct 2018

On the distribution of Samotherium major: New data from the Turolian of Greece and Turkey

Nagel, D. (Contributor), Xafis, A. (Speaker), Grimsson, F. (Contributor), Mayda, S. (Contributor), Kaya, T. (Contributor) & Halaçlar, K. (Contributor)

9 Oct 2018

Funktionsmorphologische Anpassungen des knöchernen Labyrinthes bei Beuteltieren.

Pfaff, C. (Speaker), Czerny, S. (Contributor), Nagel, D. (Contributor) & Kriwet, J. (Contributor)

24 Mar 2017 → 26 Mar 2017

Die fossilen Carnivoren der Hominidenfundstelle Galili (Äthiopien)

Nagel, D. (Speaker)

7 Oct 2009

Curriculum Vite

1988 Assistant Research Scientist: Austrian Science Foundation, Project No. 6514 E: Evolution and Chronology of the Cave Bear.

1988-1992 Assistant Research Scientist at the Department of Paleontology of Vienna
Curator of the micro-mammal collections at the Department of Paleontology.

1992-2001 Associate Research Scientist at the Department of Paleontology, Vienna

2001 – Associated Professor at the Department of Paleontology, Vienna
Responsible for the micro-mammal collection. Editor of the paleontological journal Beiträge zur Paläontologie. Organizer of Conferences for the Mammal section of the Department. Organizing and executing excavations.

1998 Radiation Protection Training at the Austrian Research Center Seibersdorf.

2009 to 2010 Head of the Department of Palaeontology. Since 2006 responsible for teaching administration of Palaeobiology at the Life Science Faculty in Vienna.

From 2011 to 2014 again Head of the Department of Palaeontology

From 1988-1990 and 1996-1998 Secretary of the Austrian Palaeontological Society, from 2002-2004 President of the Austrian Palaeontological Society, since 2002 President of the Society Friends of Palaeontology; since 2002-2012 member of the Quaternary Commission of the Austrian Academy of Sciences. 2007-2009 on Member-at-Large of the Palaeontological Society (Germany), 2007 - 2012 member of the Gregory Award Commitee, Society of Vertebrate Paleontology. 2011-2014 Vice-President of the Palaeontological Association (Germany); 2011-2013 Member of the Ethics Commission at the University of Vienna.

Als es der Hyäne bei uns zu kalt wurde. 50.000 Jahre Klimageschichte aus der Teufelslucke bei Roggendorf.

Nagel, D. (Speaker)

28 Oct 2022

Activity: Talks and presentations › Talk or oral contribution › Science to Public

Die Teufelslucke bei Eggenburg – Klimaarchiv der letzten 50.000 Jahre

Nagel, D. (Speaker)

4 Jul 2022

Activity: Talks and presentations › Talk or oral contribution › Science to Public

Pint of Science- tales from the 50 million year old amber forest

Seyfullah, L. J. (Speaker) & Nagel, D. (Speaker)

11 May 2022

Activity: Talks and presentations › Talk or oral contribution › Science to Public